Genus Hexatylus
Goodey, 1926
Syn.:
|
Anguillulina (Hexatylus Goodey, 1926) (Schneider
1939)
Neotylenchus Steiner, 1931
Anguillulina (Neotylenchus Steiner, 1931)
(Schneider 1939)
Scytaleum Andrássy, 1961 |
Diagnosis
Neotylenchinae. Entomoparasitic generation:
Adult female in insect haemocoel, elongate-obese. Males and preadult
infective female resembling those of most Sphaerularioidea, female with
much larger stylet bearing basal knobs and having enlarged oesophageal
glands, extremely elongate uterus filled with minute sperm and immature
ovary. Preadult female penetrates an insect host and matures in its
haemocoel. Free-living, mycetophagous generation: Female:
Body veriform, straight or slightly arcuate ventrally upon relaxation,
about 1 mm long (0.76 - 0.9 mm long, maximum width 45 - 57 um in
type species). Cuticle finely striated. Lateral fields each
with six incisures, not areolated. Cephalic region very low, continuous
or slightly offset, smooth; labial disc absent; oral opening circular,
six inner labial and four cephalic papillae present; amphid apertures small,
labial but slightly posterior to level of cephalic papillae; framework
six-sectored at base, 12-sectored anteriorly. Stylet slender, about
8 -12 um long; conus shorter than shaft; basal knobs in type species
duplex or bifid so that they appear as six swelllings; two to three guidings
rings present around shaft. Some stylet protractors pass through
framework to attach to cuticle on top of head. Oesophagus cylindroid
or fusiform. Nerve ring circum-intestinal. Excretory pore
behind nerve ring. Oesophageal glands overlapping intestine, dorsal
gland larger that subventrals. Vulva a long transverse slit, usually
at less than two body-widths from anus. Vagina directed anteriorly.
No postvulval uterine sac. Prodelphic. Uterus with a prominent
crustaformeria. Eggs elongate-oval; one or two eggs in uterus at
any one time. Oviparous. Oviduct with two consecutive rings
of four cells each or two rows of seven to nine cells (Geraert, 1976).
Ovary outstretched, tip may be reflexed. Female tail elongate-conoid,
pointed, may be filiform. Rectum and anus distinct; rectum acts
as a feeding pump. Male: Stylet present. Testis producing
minute rounded sperm. Spicules anteriorly expanded, arcuate, cephalated.
Gubernaculum simple, fixed. Bursa distinct, terminal or subterminal.
Juveniles essentially similar to female.
(Description- Siddiqi, 2000)
Type species
Hexatylus viviparus Goodey, 1926
syn.: Anguillulina (Hexatylus)
vivipara
(Goodey)
W.Schneider, 1939
Iotonchium
vivaparum (Goodey) Filipjev & Schuurmans Stekhoven, 1941
Neotylenchus
abulbosus Steiner, 1931
Anguillulina
abulbosus (Steiner) Schneider, 1939
Hexatylus
abulbosus (Steiner) Goodey, 1933
Iotonchium
vivaparum (Steiner) Filipjev & Schuurmans Stekhoven, 1941
Other Species
Hexatylus
acutus (Thorne, 1941) Siddiqi, 1986
syn.: Neotylenchus acutus Thorne, 1941 (N. apiculatus in
Thorne, 1941 = lapsus calami)
H.
coprophagus Goodey, 1938
syn.: Iotonchium coprophagum (Goodey) Filipjev & Schuurmans
Stekhoven, 1941
Neotylenchus coprophagus (Goodey) Thorne, 1941
H.
kirjanovae (Izatullaeva, 1969) Siddiqi, 1986
syn.: Neotylenchus kirjanovae Izatullaeva, 1969
H.
mucronatus (Izatullaeva, 1969) Siddiqi, 1986
syn.: Neotylenchus mucronatus Izatullaeva, 1969
H.
mulveyi Das, 1964
H.
nudus (Thorne & Malek, 1968) Siddiqi, 1986
syn.: Neotylenchus nudus Thorne & Malek, 1968
H.
vigissi Skarbilovich, 1952
syn.: Neotylenchus vigissi (Skarbilovich) Goodey, 1963
Scytaleum vigissi (Skarbilovich) Andrássy, 1961
DNA Sequences Obtained